Actually, it turns out that the ferrungulate tarsus is a fairly apt study for the understanding of ankles, assuming that one is apt to study such things at all.
In addition to the pretarsus, tarsus , tibia and femur, insects have two proximal (closer to the body) segments not shown in the figure.
Preserved bones of the tarsus include the astragalus, calcaneum, and medial centrale.
A cockroach leg is made up of a number of distinct segments, arranged along the proximo-distal axis in the order coxa, femur, tibia, tarsus .
Both sexes have a pair of ‘spurs’ on each hind leg where the tibia, or fourth leg segment, joins the tarsus , or foot.
A wedge of skin, orbicularis, and tarsus is removed parallel to the lid margin.
An influential contribution is that of Szalay, who linked Australian taxa to one group among the South American radiation, based on structure of the tarsus .
From 1983 to 1997, trapping teams captured adults at nest sites and banded each unmarked bird with a numbered band on the right tarsus and an alpha-numeric color band on the left tarsus .
Tarsus color turns black to red between yearling and adult, which suggests that the appearance of the tarsus could be involved in signal of status.
As with enlarged leg musculature, larger tarsi may be linked to improved adeptness at locomotion for more effective foraging and predator evasion.